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Tangkoko forest reserve, North Sulawesi, Indonesia. 14-03-14

Tangkoko forest reserve, North Sulawesi, Indonesia. 14-03-14

14/03/14 Tangkoko forest, North Sulawesi, Indonesia

Tangkoko is one of the last remaining strongholds of the Critically Endangered Sulawesi crested black macaques (Macaca nigra). Wildlife photographer Andrew Walmsley is in the field documenting the work of Selamatkan Yaki, a non-governmental organisation (NGO) working to protect the monkeys which are threatened by habitat loss and the illegal bushmeat trade.

A female macaque snaps at a male just out of shot- baring their teeth is a threat display, and scuffles like this are common in macaque communities.

To take these photos, Andrew sat at the top of the tree from just after 5am until after 1pm.